Skip to Content
Sasha Reed

Sasha Reed

Research Ecologist, Southwest Biological Science Center, U.S. Geological Survey

Southwest Biological Science Center
U.S. Geological Survey
2290 S. West Resource Blvd
Moab, UT 84532


  • Research Ecologist, Southwest Biological Science Center, U.S. Geological Survey


  • PhD, Biogeochemistry, University of Colorado-Boulder, 2008
  • BA, Organic Chemistry, Colgate College, 1997

External Links

Journal Articles

In Press

Faist, A. M., A. J. Antoninka, J. Belnap, M. A. Bowker, M. C. Duniway, F. Garcia-Pichel, C. Nelson, S. C. Reed, A. Giraldo-Silva, S. Velasco-Ayuso and N. N. Barger. Inoculation and habitat amelioration efforts in biological soil crust recovery vary by desert and soil texture. Restoration Ecology DOI: 10.1111/rec.13087. (link )


Bachelot, B., A. M. Alonso-Rodriguez, L. Aldrich-Wolfe, M. A. Cavaleri, S. C. Reed and T. E. Wood. 2020. Altered climate leads to positive density‐dependent feedbacks in a tropical wet forest. Global Change Biology 26(6):3417-3428. DOI: 10.1111/gcb.15087. (link )

Chuckran, P. F., R. Reibold, H. L. Throop and S. C. Reed. 2020. Multiple mechanisms determine the effect of warming on plant litter decomposition in a dryland. Soil Biology and Biochemistry 145(Jun):107799. DOI: 10.1016/j.soilbio.2020.107799. (link )

Delgado-Baquerizo, M., P. B. Reich, C. Trivedi, D. J. Eldridge, S. Abades, F. D. Alfaro, F. Bastida, A. A. Berhe, N. A. Cutler, A. Gallardo, L. García-Velázquez, S. C. Hart, P. E. Hayes, J. He, Z. Hseu, H. Hu, M. Kirchmair, S. Neuhauser, C. A. Perez, S. C. Reed, F. Santos, B. W. Sullivan, P. Trivedi, J. Wang, L. Weber-Grullon, M. A. Williams and B. K. Singh. 2020. Multiple elements of soil biodiversity drive ecosystem functions across biomes. Nature Ecology & Evolution 4:210-220. DOI: 10.1038/s41559-019-1084-y. (link )

Hoover, D. L., B. Bestelmeyer, N. B. Grimm, T. E. Huxman, S. C. Reed, O. E. Sala, T. R. Seastedt, H. Wilmer and S. Ferrenberg. 2020. Traversing the wasteland: A framework for assessing ecological threats to drylands. BioScience 70(1):35-47. DOI: 10.1093/biosci/biz126. (link )


Delgado-Baquerizo, M., R. D. Bardgett, P. M. Vitousek, F. Maestre, M. A. Williams, D. J. Eldridge, H. Lambers, S. Neuhauser, A. Gallardo, L. García-Velázquez, O. E. Sala, S. Abades, F. D. Alfaro, A. A. Berhe, M. A. Bowker, C. M. Currier, N. A. Cutler, S. C. Hart, P. E. Hayes, Z. Hseu, M. Kirchmair, V. M. Pena-Ramirez, C. A. Perez, S. C. Reed, F. Santos, C. Siebe, B. W. Sullivan, L. Weber-Grullon and N. Fierer. 2019. Changes in belowground biodiversity during ecosystem development. Proceedings of the National Academy of Sciences of the United States of America 116(14):6891-6896. DOI: 10.1073/pnas.1818400116. (link )


Antoninka, A., R. W. Bowker, P. Chuckran, N. N. Barger, S. C. Reed and J. C. Belnap. 2018. Maximizing establishment and survivorship of field-collected and greenhouse-cultivated biocrusts in a semi-cold desert. Plant and Soil 429(1-2):213-225. DOI: 10.1007/s11104-017-3300-3. DOI: 1573-5036. (link )

Bowker, M. A., S. C. Reed, F. Maestre and D. J. Eldridge. 2018. Biocrusts: The living skin of the earth. Plant and Soil 429(1-2):1-7. DOI: 10.1007/s11104-018-3735-1. (link )

Darrouzet-Nardi, A., S. C. Reed, E. E. Grote and J. C. Belnap. 2018. Patterns of longer-term climate change effects on CO2 efflux from biocrusted soils differ from those observed in the short term. Biogeosciences 15(14):4561-4573. DOI: 10.5194/bg-15-4561-2018. (link )

Feng, X., M. Uriarte, G. Gonzalez, S. Reed, J. Thompson, J. K. Zimmerman and L. Murphy. 2018. Improving predictions of tropical forest response to climate change through integration of field studies and ecosystem modeling. Global Change Biology 24:e213-e232. DOI: 10.1111/gcb.13863. (link )

Grossiord, C., A. Gessler, S. C. Reed, I. Borrego, A. D. Collins, L. T. Dickman, M. Ryan, L. Schönbeck, S. Sevanto, A. Vilagrosa and N. G. McDowell. 2018. Reductions in tree performance during hotter droughts are mitigated by shifts in nitrogen cycling. Plant, Cell & Environment 41(11):2627-2637. DOI: 10.1111/pce.13389. (link )

Mau, A. C., S. C. Reed, T. E. Wood and M. A. Cavaleri. 2018. Temperate and tropical forest canopies area already functioning beyond their thermal thresholds for photosynthesis. Forests 9(1):47. DOI: 10.3390/f9010047. (link )

Munson, S. M., S. C. Reed, J. Penuelas, N. G. McDowell and O. E. Sala. 2018. Ecosystem thresholds, tipping points, and critical transitions. New Phytologist Trust 218(4):1315-1317. DOI: 10.1111/nph.15145. (link )

Torres-Cruz, T. J., A. J. Howell, R. H. Reibold, T. A. McHugh, M. A. Eickhoff and S. C. Reed. 2018. Species-specific nitrogenase activity in lichen-dominated biological soil crusts from the Colorado Plateau, USA. Plant and Soil 429(1-2):113-125. DOI: 10.1007/s11104-018-3580-2. (link )

Tucker, C. L., S. Ferrenberg and S. C. Reed. 2018. Climatic sensitivity of dryland soil CO2 fluxes differs dramatically with biological soil crust successional state. Ecosystems 1-18. DOI: 10.1007/s10021-018-0250-4. (link )

Tucker, C., D. Yan, M. Dannenberg, S. C. Reed and W. Smith. 2018. Science at the frontier: Multimethod research to evaluate ecosystem change across multiple scales. New Phytologist 218(4):13118-1320. DOI: 10.1111/nph.15195. (link )

Vicca, S., B. D. Stocker, S. C. Reed, W. R. Wieder, M. Bahn, P. A. Fay, I. A. Janssens, H. Lambers, J. Penuelas, S. Piao, K. T. Rebel, J. Sardans, B. D. Sigurdsson, K. Van Sundert, Y. Wang, S. Zaehle and P. Ciais. 2018. Using research networks to create the comprehensive datasets needed to assess nutrient availability as a key determinant of terrestrial carbon cycling. Environmental Research Letters 13(12):125006. DOI: 10.1088/1748-9326/aaeae7. (link )

Wertin, T. M., K. E. Young and S. C. Reed. 2018. Spatially explicit patterns in a dryland's soil respiration and relationships with climate, whole plant photosynthesis and soil fertility. Oikos 127(9):1280-1290. DOI: 10.1111/oik.04935 . (link )

Winkler, D. E., D. Backer, J. C. Belnap, J. B. Bradford, B. Butterfield, S. M. Copeland, M. C. Duniway, A. M. Faist, S. E. Fick, S. L. Jensen, A. T. Kramer, R. Mann, R. T. Massatti, M. L. McCormick, S. M. Munson, P. Olwell, S. D. Parr, A. A. Pfennigwerth, A. M. Pilmanis, B. A. Richardson, E. Samuel, K. See, K. E. Young and S. C. Reed. 2018. Beyond traditional ecological restoration on the Colorado Plateau. Restoration Ecology 26(6):1055-1060. DOI: 10.1111/rec.12876. (link )


Chung, Y. A., R. L. Sinsabaugh, C. R. Kuske, S. C. Reed and J. A. Rudgers. 2017. Spatial variation in edaphic characteristics is a stronger control than nitrogen inputs in regulating soil microbial effects on a desert grass. Journal of Arid Environments 147(Jul):59-65. DOI: 10.1016/j.jaridenv.2017.03.005. (link )

Ferrenberg, S., A. M. Faist, A. J. Howell and S. C. Reed. 2017. Biocrusts enhance soil fertility and Bromus tectorum growth, and interact with warming to influence germination. Plant and Soil 429(1-2):77-90. DOI: 10.1007/s11104-017-3525-1. (link )

Ferrenberg, S. and S. C. Reed. 2017. Biocrust ecology: Unifying micro- and macro-scales to confront global change. New Phytologist 216(3):643-646. DOI: 10.1111/nph.14826. (link )

Ferrenberg, S., C. L. Tucker and S. C. Reed. 2017. Biological soil crusts diminutive communities of potential global importance. Frontieres in Ecology and the Environment 15(3):160-167. DOI: 10.1002/fee.1469. (link )

Lu, X., S. C. Reed, S. Hou, Y. Hu, H. Wei, F. Lu, Q. Cui and X. Han. 2017. Temporal variability of foliar nutrients: Responses to nitrogen deposition and prescribed fire in a temperate steppe. Biogeochemistry 133(3):295-305. DOI: 10.1007/s10533-017-0333-x. (link )

McHugh, T. A., E. M. Morrissey, R. C. Mueller, L. Gallegos-Graves, C. R. Kuske and S. C. Reed. 2017. Bacterial, fungal, and plant communities exhibit no biomass or compositional response to two years of simulated nitrogen deposition in a semiarid grassland. Environmental Microbiology 19(4):1600-1611. DOI: 10.1111/1462-2920.13678. (link )

Rutherford, W. A., T. H. Painter, S. Ferrenberg, J. Belnap, G. S. Okin, C. Flagg and S. C. Reed. 2017. Albedo feedbacks to future climate via climate change impacts on dryland biocrusts. Scientific Reports 7:Art. 44188. DOI: 10.1038/srep44188. (link )

Tucker, C. L., T. A. McHugh, A. Howell, R. Gill, B. Weber, J. C. Belnap, E. Grote and S. C. Reed. 2017. The concurrent use of novel soil surface microclimate measurements to evaluate CO2 pulses in biocrusted interspaces in a cool desert ecosystem. Biogeochemistry 135(3):239-249. DOI: 10.1007/s10533-017-0372-3. (link )

Wertin, T. M., J. Belnap and S. C. Reed. 2017. Experimental warming in a dryland community reduced plant photosynthesis and soil CO2 efflux although the relationship between the fluxes remained unchanged. Functional Ecology 31(2):297-305. DOI: 10.1111/1365-2435.12708. (link )

Whitney, K. M., E. R. Vivoni, M. C. Duniway, J. B. Bradford, S. C. Reed and J. C. Belnap. 2017. Ecohydrological role of biological soil crusts across a gradient in levels of development. Ecohydrology 10(7):e1875. DOI: 10.1002/eco.1875. (link )

Young, K. E. and S. C. Reed. 2017. Spectrally monitoring the response of the biocrust moss Syntrichia caninervis to altered precipitation regimes. Scientific Reports 7:Art. 41793. DOI: 10.1038/srep41793. (link )

Book Chapters


Steven, B., T. A. McHugh and S. C. Reed. 2017. 8. The response of arid soil communities to climate change. Pp. 139-158 In: Steven, B. ed., The Biology of Arid Soils. De Gruyter. ISBN: 978-3-11-041904-7. (link )



Whitney, K. M., E. R. Vivoni, M. C. Duniway, J. B. Bradford, S. C. Reed and J. C. Belnap. 2015. Role of biological soil crusts in hydrologic modeling of western high-desert ecosystems.. Poster presented at the Third Annual Arizona Geological Society Doug Shakel Memorial Student Poster Event, Tempe, AZ.



Whitney, K. M., E. R. Vivoni, M. C. Duniway, J. B. Bradford, S. C. Reed and J. C. Belnap. 2015. Role of biological soil crusts in hydrologic modeling of western high-desert ecosystems. Presentation at the American Meteorological Society Annual Meeting, Phoenix, AZ.